Wilhelmina Slater is a diva and former supermodel who works as the Creative Director of Mode.  However, she will do whatever it takes to get the role of Editor in Chief that she feels was unfairly given to Daniel.  She does claim it for a bit in season 3 but loses the title again by the end of episode 2.  She is now co-Editor in Chief alongside Daniel.

Wilhelmina Slater Quotes

Who cares what the average woman wears on the street. We shouldn't be taking pictures of them, we should be throwing rocks at them.

Wilhelmnia

The rounded o's of a stunted adolescent. The stiff t's of someone who is clearing overcompensating. This is Daniel's handwriting.

Wilhelmnia
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Ugly Betty Quotes

Hilda: The test just said I had to do a manicure, it didn't say on who.
[Hilda shows off her nails]
Hilda: I wasn't gonna waste all this hotness on someone else

I smell a burrito

Wilhelmina [holding up Betty's earring]